Revisionary studies of the mayfly family Baetidae (Ephemeroptera)
Abstract
Two previously unrecognized and monophyletic generic complexes of Baetidae are identified on the basis of larval and adult character states including (1) the Cloeon complex of genera (Baetopus Keffermuller, Centroptilum Eaton, Cloeon Leach, Neocloeon Traver, Procloeon Bengtsson, Raptobaetopus Muller-Liebenau, and possibly, Pseudocentroptilum Bogoescu) and (2) the Baetis complex of genera (Acentrella Bengtsson, Baetiella Ueno, Baetis Leach, Barbaetis Waltz and McCafferty, Heterocloeon McDunnough, Jubabaetis Muller-Liebenau, Liebebiella gen. n, Platybaetis Muller-Liebenau, and Pseudocloeon Klapalek). Within the Cloeon complex of genera Procloeon is revised and shown to include many Nearctic species previously placed in Centroptilum. Centroptilum is restricted to include only its type species. Neocloeon is reinstated as a valid taxon and revised. Within the Baetis complex of genera, Pseudocloeon is restricted to include only its type species from Java. Oriental larvae provisionally classified in Pseudocloeon are placed in a new genus Liebebiella gen. n. Other taxonomic revisions not associated with either of the two monophyletic complexes above include the following. The Neotropical genus Moribaetis gen. n. is described to include two nominal subgenera Mayobaetis subgen. n. and Moribaetis s.s. Three Afrotropical genera are described, Acanthiops gen. n., Pseudopannota gen. n., and Ophelmastoma gen. n., based on highly atypic larval characters. Cloeodes Traver (=Centroptella Braasch and Soldan, in part syn.n.) is revised to include two subgenera, Notobaetis Morihara and Edmunds stat. n., and Cloeodes s.s. As redefined Cloeodes includes both Western Hemisphere and Oriental species. A sister genus of Cloeodes from South America is identified, Bernerius gen.n. and a South American origin of the genus is hypothesized. Larvae previously placed in Centroptella are shown to be in part assignable to Cloeodes s.s. including the previously declared type of Centroptella, other species are placed in a new genus Chopralla gen. n.
Degree
Ph.D.
Advisors
McCafferty, Purdue University.
Subject Area
Entomology
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